Certain shading techniques now seem suggested by the the subject, for example, hatching for the markings on the wren's tail feathers or stippling on the cattail reed. The negative space concept (ingrained by the upside down drawing) was very useful in getting proportion and shape of the wren to reed areas. My biggest downfall is with rapid gesture drawing (my third attempt at a one minute wren yielded woefully inaccurate proportions). There seems to be inadequate time to hold out the pencil to measure and still get the shapes and essence down. Hopefully it will improve with practice. In my refined drawing the wren appears to be more in profile than the photo depicts. Perhaps a problem with the eye position or shape or the beak or neck shading...?
1. Drawing from the photo gave me time to look many times at proportions and details of a perfectly posed subject. Nothing came easy, but the proportions were very difficult and the expression on this cute warbler's face was very difficult to capture.
2. I would not have noticed the way the beak meets the head, the detail of the feet gripping the branch with the long "fingers" extending below the branch, the wing coverts and many other details. Nature journaling gets one to observe these details, but, if one is drawing a warbler in nature and not from a photo, the observation time will be too short for me to get that sort of detail.